Rare Middle and Upper Devonian dalmanelloid (Orthida) of the Cantabrian Mountains, N Spain

Abstract

Rare Cantabrian Dalmanellidae (Costisorthis lisae nov. sp.), Dicoelosiidae (Teichertina cf. peregrina, T. cf. ?tzroyensis), and Mystrophoridae (Mystrophora sp., Biernatium sucoi nov. sp., and Biernatium sp. 2) are described and ?gured for the ? rst time in Spain. Most of them are scarce forms that occur only in certain localities. All the species but one came from the lower part of the Portilla (province of León) and Candás (province of Asturias) formations, Faunal Interval 21, Polygnathus rhenanus/P. varcus conodont zone, middle Givetian. The exception is Biernatium sp. 2 that occurs in Asturias in FI 25, Piñeres Fm., Palmatolepis transitans zone, lower Frasnian. Costisorthis ranges from Pragian to Eifelian rocks in central Europe. C. lisae nov. sp., from the Givetian of Asturias di?ers from other Costisorthis species in the weaker development of the distinctive ventral and dorsal plications.